@layer reset, tokens, layout, components, pages, responsive;
@layer reset {
  *,*::before,*::after{box-sizing:border-box}
  body,h1,h2,h3,p,figure,dl,dd{margin:0}
  img,svg{max-width:100%;display:block}
  input,button,select,textarea{font:inherit}
  button,summary{cursor:pointer}
  a{color:inherit}
  [hidden]{display:none!important}
}
@layer tokens {
  :root{color-scheme:light;--paper:#f5f5eb;--surface:#fffefa;--surface-alt:#edf0e5;--text:#182a24;--muted:#4c6258;--line:#dce4d7;--forest:#164e42;--forest-dark:#0c342b;--mint:#d7ecb2;--lime:#d9efad;--bad:#94352a;--shadow:0 8px 24px #142a2020;--radius:18px;--display:Georgia,"Times New Roman",serif;--ui:system-ui,-apple-system,BlinkMacSystemFont,"Segoe UI",sans-serif}
  :root[data-theme=dark]{color-scheme:dark;--paper:#101c18;--surface:#192a23;--surface-alt:#20362b;--text:#eef5e8;--muted:#b6c7bb;--line:#3b5247;--forest:#c4e4ad;--forest-dark:#102f26;--mint:#d7ecb2;--bad:#ffb3a9;--shadow:0 8px 24px #0004}
}
@layer layout {
  html{scroll-padding-top:88px}
  body{background:var(--paper);color:var(--text);font:16px/1.62 var(--ui);min-width:280px}
  main{min-height:65vh}
  .shell{width:min(calc(100% - 72px),1320px);margin-inline:auto}
  .page{padding-block:52px 72px}
  .home{padding-top:25px;padding-bottom:64px}
  .narrow{max-width:780px}
  h1,h2,h3{font-family:var(--display);line-height:1.05;font-weight:500;letter-spacing:-.025em;text-wrap:balance}
  h1{font-size:clamp(2.75rem,6.1vw,5.1rem);margin:.12em 0 .22em}
  h2{font-size:clamp(1.95rem,3.5vw,3.25rem);margin:.12em 0 .33em}
  h3{font-size:1.53rem;margin-bottom:.18em}
  p+p{margin-top:.75em}
  section p+form,section p+.actions,form+details{margin-top:16px}
  section[id],form[id]{scroll-margin-top:100px}
  .site-header{position:sticky;top:0;z-index:10;background:var(--surface);border-bottom:1px solid var(--line)}
  .header-row{height:70px;display:flex;align-items:center;justify-content:space-between;gap:22px}
  .navigation,.tools,.brand,.footer-row{display:flex;align-items:center;gap:20px}
  .brand{font:600 1.32rem/1 var(--display);text-decoration:none;letter-spacing:-.025em;gap:9px;white-space:nowrap}
  .navigation a{position:relative;font-weight:600;font-size:.91rem;text-decoration:none;padding:22px 5px;color:var(--muted)}
  .navigation a[aria-current]{color:var(--text)}
  .navigation a[aria-current]::after{content:"";position:absolute;inset:auto 5px 0;height:3px;background:var(--forest);border-radius:4px}
  .tools{gap:8px}
  .site-footer{border-top:1px solid var(--line);background:var(--surface);padding-block:28px;color:var(--muted)}
  .footer-row{justify-content:space-between;flex-wrap:wrap;font-size:.9rem}
  .footer-row .brand{color:var(--text)}
  .skip,.visually-hidden{position:absolute;width:1px;height:1px;overflow:hidden;clip-path:inset(50%)}
  .skip:focus{clip-path:none;width:auto;height:auto;top:8px;left:10px;z-index:50;background:var(--surface);padding:8px 14px}
  :focus-visible{outline:3px solid #a57830;outline-offset:4px}
}
@layer components {
  .eyebrow{font-size:.73rem;font-weight:760;letter-spacing:.16em;text-transform:uppercase;color:var(--forest);line-height:1.4}
  .quiet,.scientific,small{color:var(--muted)}
  .intro{max-width:66ch;font-size:1.12rem;color:var(--muted);margin-bottom:22px}
  .button{display:inline-flex;align-items:center;justify-content:center;gap:10px;min-height:43px;padding:8px 18px;border:1px solid transparent;border-radius:9px;background:var(--forest);color:var(--surface);font-size:.9rem;font-weight:700;text-decoration:none;line-height:1.35}
  .button:hover{filter:brightness(1.13)}
  .button.outline{background:transparent;border-color:var(--line);color:var(--text)}
  .button.primary{background:var(--lime);color:#16281a}
  .button.ghost{background:#173e31b0;border-color:#d7ecb255;color:#fcfff8}
  .button.compact{min-height:36px;padding:6px 12px}
  .actions,.input-row,.tab-links{display:flex;align-items:center;gap:10px;flex-wrap:wrap}
  .text-link,.text-button{color:var(--forest);font-weight:700;font-size:.92rem}
  .text-button{border:0;background:transparent;text-decoration:underline;padding:3px 4px}
  .icon-button{width:40px;height:40px;display:grid;place-items:center;border:1px solid var(--line);border-radius:10px;background:var(--surface);color:var(--text);list-style:none}
  summary::-webkit-details-marker{display:none}
  .icon-button:hover{background:var(--surface-alt)}
  .icon-button svg{width:19px;height:19px;stroke:currentColor;stroke-width:1.65;fill:none;stroke-linecap:round;stroke-linejoin:round}
  .menu{position:relative}
  .popover{position:absolute;right:0;top:calc(100% + 10px);width:min(360px,calc(100vw - 24px));padding:16px;box-shadow:var(--shadow);border-radius:var(--radius);border:1px solid var(--line);background:var(--surface)}
  .search-menu .popover{right:-96px}
  .more{display:grid;gap:10px;width:230px}
  .more a{font-weight:600;text-decoration:none}
  .more .mobile-only{display:none}
  .more .quiet{border-top:1px solid var(--line);padding-top:8px;font-size:.85rem}
  .quick-search>label{display:block;margin-bottom:6px}
  .quick-search input,.input-row input{min-width:0;flex:1}
  .suggestions,.plain-list{list-style:none;padding:0;margin:12px 0 0}
  .suggestions a{display:flex;align-items:center;gap:9px;padding:7px;border-radius:8px;text-decoration:none}
  .suggestions a:hover{background:var(--surface-alt)}
  .suggestions img{width:38px;height:38px;object-fit:cover;border-radius:7px}
  .suggestions span{display:grid;line-height:1.35}
  .errorlist{color:var(--bad)}
  .toast-stack{position:fixed;z-index:40;right:max(12px,env(safe-area-inset-right));bottom:max(12px,env(safe-area-inset-bottom));width:min(360px,calc(100% - 32px));max-height:50vh;overflow:auto;display:grid;gap:8px;pointer-events:none}
  .toast{display:flex;align-items:center;gap:10px;padding:8px 12px;border:1px solid var(--line);border-inline-start:4px solid var(--forest);border-radius:10px;background:var(--surface);color:var(--text);box-shadow:var(--shadow);pointer-events:auto}
  .toast.error{border-inline-start-color:var(--bad)}
  .toast p{flex:1;margin:0;overflow-wrap:anywhere}
  .toast button{flex:none;min-width:36px;min-height:36px;border:0;border-radius:7px;background:var(--surface-alt);color:var(--text);font-size:1.2rem;cursor:pointer}
  .panel{padding:clamp(16px,3vw,28px);background:var(--surface);border:1px solid var(--line);border-radius:var(--radius);min-width:0}
  .content-section,.home-section{margin-top:clamp(34px,6vw,70px)}
  .section-heading{display:flex;align-items:end;justify-content:space-between;gap:16px;margin-bottom:15px}
  .section-heading h1,.section-heading h2{margin-bottom:0}
  .card-grid,.branch-grid,.fact-grid{display:grid;grid-template-columns:repeat(auto-fit,minmax(min(100%,235px),1fr));gap:15px}
  .card-grid{grid-template-columns:repeat(4,minmax(0,1fr))}
  .organism-card{display:flex;flex-direction:column;overflow:hidden;border:1px solid var(--line);border-radius:var(--radius);background:var(--surface);min-width:0}
  .organism-card h3 a{text-decoration:none}
  .organism-card h3 a:hover{text-decoration:underline}
  .card-visual{aspect-ratio:1.55;background:#d6e0c8;overflow:hidden;display:grid;place-items:center}
  .card-visual img{width:100%;height:100%;object-fit:cover;transition:transform .25s ease}
  .organism-card:hover .card-visual img{transform:scale(1.045)}
  .card-visual.abstract{background:radial-gradient(ellipse at 70% 0,#536f50,#1b5145 60%,#123a31);color:#d9efad90}
  .abstract svg{width:65px;fill:none;stroke:currentColor;stroke-width:.7}
  .card-copy{padding:15px 17px 17px}
  .card-summary{font-size:.87rem;margin-top:10px;color:var(--muted)}
  .scientific{font-family:var(--display);margin-top:4px}
  .card-list{display:grid;gap:12px}
  .card-list .organism-card{display:grid;grid-template-columns:145px minmax(0,1fr)}
  .card-list .card-visual{height:100%;aspect-ratio:auto}
  .branch-card{display:grid;gap:5px;padding:17px 19px;min-height:100px;border-radius:13px;border:1px solid var(--line);background:var(--surface);text-decoration:none}
  .branch-card:hover{background:var(--surface-alt)}
  .branch-card strong{font:1.48rem var(--display)}
  .branch-card>span:last-child:not(.eyebrow){justify-self:end;color:var(--forest)}
  .fact-grid>div{background:var(--surface);padding:14px 17px;border:1px solid var(--line);border-radius:12px}
  .fact-grid dt{font-size:.83rem;color:var(--muted);font-weight:700}
  .fact-grid dd{font:1.45rem/1.2 var(--display);margin:.2em 0}
  .plain-list li{padding:8px 0;border-bottom:1px solid var(--line)}
  .table-wrap{overflow:auto;border:1px solid var(--line);border-radius:12px;background:var(--surface)}
  table{width:100%;border-collapse:collapse;text-align:left}
  th,td{padding:11px 14px;border-bottom:1px solid var(--line);vertical-align:top}
  thead th{background:var(--surface-alt)}
  tr:last-child td,tr:last-child th{border-bottom:0}
  label{display:grid;gap:5px;font-weight:650;color:var(--text);font-size:.9rem}
  input:not([type=radio]):not([type=checkbox]),select,textarea{max-width:100%;color:var(--text);background:var(--surface);border:1px solid var(--line);border-radius:8px;padding:8px 10px;min-height:42px}
  select{padding-right:18px}
  textarea{width:100%;resize:vertical}
  .form-stack{display:grid;gap:12px;align-content:start}
  .form-stack p{margin:0;display:grid;gap:5px}
  .form-stack .button{justify-self:start}
  .helptext{font-size:.82rem;color:var(--muted);overflow-wrap:anywhere}
  .filters{display:flex;align-items:end;flex-wrap:wrap;gap:12px;margin-bottom:20px}
  .filters label:first-child{flex:1;min-width:min(230px,100%)}
}
@layer pages {
  .hero{position:relative;isolation:isolate;min-height:500px;border-radius:23px;overflow:hidden;background:#10382e;color:#fff;display:grid;align-items:center;box-shadow:var(--shadow)}
  .hero::after{content:"";position:absolute;z-index:-1;inset:0;background:linear-gradient(90deg,#0d3127 5%,#0d3127 34%,#0d3127d5 45%,#0d312725 70%,#0d312708);pointer-events:none}
  .hero-photo{position:absolute;inset:0 0 0 auto;height:100%;width:66%;object-fit:cover;object-position:center 39%;z-index:-2}
  .hero-copy{padding:55px clamp(28px,6vw,78px);max-width:720px}
  .hero-copy h1{font-size:clamp(3.2rem,7.3vw,6.45rem);letter-spacing:-.065em;margin:.14em 0 .18em;line-height:.94}
  .hero-copy h1 em{font-weight:400;color:#d9efad}
  .hero .eyebrow,.learning .eyebrow{color:#d9efad;display:flex;align-items:center;gap:7px}
  .hero-copy>p:not(.eyebrow){color:#f0f4e7;max-width:39ch;font-size:1.06rem}
  .hero-copy .actions{margin-top:24px}
  .dot{width:7px;height:7px;border-radius:50%;background:#d9efad}
  .hero-credit{position:absolute;bottom:14px;right:15px;padding:4px 9px;background:#112218c7;border-radius:6px;color:white;font-size:.71rem}
  .learning{background:#143c32;color:white;border-radius:21px;padding:32px clamp(22px,5vw,60px);display:grid;grid-template-columns:1.25fr .85fr;align-items:center;gap:25px}
  .learning .button{margin-top:17px}
  .learning-links{display:grid;gap:2px}
  .learning-links a{display:flex;justify-content:space-between;gap:7px;border-bottom:1px solid #d9efad50;padding:10px 0;font-weight:650;text-decoration:none}
  .breadcrumbs{display:flex;gap:8px;flex-wrap:wrap;font-size:.86rem;color:var(--muted);margin-bottom:17px}
  .species-intro,.compare-cards,.reading-grid,.explorer{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:18px}
  .species-intro{gap:32px;align-items:center}
  .binomial{font:1.48rem var(--display);margin-bottom:12px;color:var(--forest)}
  .binomial small{font:.82rem var(--ui)}
  .portrait img,.gallery img{width:100%;height:auto;max-height:520px;object-fit:cover;border-radius:14px}
  figcaption{font-size:.78rem;color:var(--muted);margin-top:6px}
  .gallery{display:grid;grid-template-columns:repeat(4,minmax(0,1fr));gap:10px;margin-top:14px}
  .blank-portrait{min-height:260px;background:var(--surface-alt);border:1px dashed var(--line);border-radius:var(--radius);display:grid;place-items:center;color:var(--muted)}
  .lineage{display:flex;flex-wrap:wrap;gap:8px;list-style:none;padding:0;margin:0 0 13px}
  .lineage a{background:var(--surface);border:1px solid var(--line);border-radius:8px;display:grid;gap:2px;padding:6px 9px;font-size:.86rem;text-decoration:none}
  .lineage a:hover{background:var(--surface-alt)}
  .lineage span{font-size:.68rem;color:var(--muted)}
  .explorer{grid-template-columns:1.65fr 1fr;align-items:start}
  .tree{list-style:none;margin:0;padding-left:17px;border-left:1px solid var(--line)}
  .tree-panel>.tree{padding:0;border:0}
  .tree summary,.tree a{padding:5px 0}
  .tree p{font-size:.82rem;margin:0 0 5px 18px}
  .tree small{padding-left:6px}
  .compare-form{display:grid;grid-template-columns:1fr 1fr auto;gap:15px;align-items:end;margin-bottom:18px}
  .compare-cards .card-visual{max-height:270px}
  .edge-list{display:grid;gap:11px}
  .edge{display:flex;align-items:center;gap:11px;flex-wrap:wrap}
  .edge p{flex:1 0 100%;font-size:.86rem;color:var(--muted)}
  .edge-type{background:var(--surface-alt);padding:4px 9px;border-radius:8px;font-weight:650}
  .quiz-question legend{padding:0 6px;font-weight:740}
  .answer{display:flex;gap:8px;align-items:center;padding:7px}
  .editor-grid{margin-top:16px;align-items:start}
  .tab-links{margin:14px 0;color:var(--forest);font-size:.9rem}
  .editor-thumb{display:inline-block;width:46px;height:46px;object-fit:cover;border-radius:7px;vertical-align:middle}
  .inline-form{display:inline}
  .note{border-bottom:1px solid var(--line);margin:10px 0;padding-bottom:10px}
  dialog{max-width:min(92vw,1100px);max-height:94vh;border:1px solid var(--line);border-radius:16px;background:var(--surface);color:var(--text);padding:12px}
  dialog::backdrop{background:#000b}
  dialog img{max-height:77vh;object-fit:contain;width:100%;margin-top:10px}
}
@layer responsive {
  @media(max-width:1000px){.card-grid{grid-template-columns:repeat(3,minmax(0,1fr))}.shell{width:calc(100% - 38px)}}
  @media(max-width:760px){
    .navigation{gap:9px}.navigation a{font-size:.81rem;padding-inline:2px}.header-row{gap:9px}.brand{font-size:1.13rem}
    .card-grid{grid-template-columns:repeat(2,minmax(0,1fr))}.hero{min-height:435px}
    .hero-photo{width:100%;opacity:.78}.hero::after{background:linear-gradient(90deg,#0d3127f0,#0d3127a8 55%,#0d31273d)}
    .species-intro,.explorer,.learning{grid-template-columns:1fr}.page{padding-top:33px}
  }
  @media(max-width:520px){
    .shell{width:calc(100% - 24px)}.header-row{height:59px}.navigation{display:none}
    .more .mobile-only{display:block}
    .popover{right:0}.search-menu .popover{right:-94px}.hero{min-height:425px}.hero-copy{padding:36px 21px 60px}
    .hero-copy h1{font-size:clamp(3rem,12vw,4.3rem)}.hero-photo{object-position:63% center}
    .hero-credit{font-size:.61rem;bottom:9px;right:10px}
    .home{padding-top:13px}.card-grid,.compare-cards,.reading-grid,.compare-form{grid-template-columns:1fr}
    .card-list .organism-card{grid-template-columns:95px minmax(0,1fr)}
    .section-heading{align-items:start;flex-direction:column}.gallery{grid-template-columns:repeat(2,minmax(0,1fr))}
    .footer-row{gap:9px}.filters label{width:100%}
  }
  @media(prefers-reduced-motion:reduce){*,*::before,*::after{animation-duration:.001ms!important;transition-duration:.001ms!important;scroll-behavior:auto!important}}
}